When assaulted, the owner takes up a strong position

TCD MS 10515 folio 91 verso

TCD MS 10515 folio 91 verso

The pellet constructed the little architect has often to defend its treasure. Other scarabs will attack it to carry off the spoil. When assaulted, the owner takes up a strong position on the summit of its pellet and, from this place of vantage, repells the attacks of the invader.
The battle won, it begins to roll. Now all is haste and bustle; it turns its back upon the ball, clutches it firmly with its mid and hind tarsi and pushing with fore legs against the sand, rolls the mass bodily backwards. The mode of rolling the pellet adopted by this large scarab is somewhat different to that assumed by the much smaller species observed in the Western Himalaya. In the latter species two beetles took an active part in the rolling of the pellet.