This sense of smell must be extremely acute

TCD MS 10515 folio 95 recto

TCD MS 10515 folio 95 recto

through the air then the beetle will wander aimlessly about and with much greater difficulty discover its pellet. And this sense of smell must be extremely acute; it must be exerted over long distances through the air. For during the passage of our column, within a few minutes after the mules had left their droppings in the desert the scarabs used to come buzzing round in numbers to roll away the refuse. They came direct on wing through the air and I have no doubt but that they smell the refuse from afar.
In addition They possss an accurate knowledge of direction for as they approached our column they always arrived in flight from the line of the river that guarded our flank to the North, and they always rolled their plundered pellets back