It would at first sight seem that there was no cooperation

TCD MS 10515 folio 92 recto

TCD MS 10515 folio 92 recto

In this species, although the pellet belongs to a pair of scarabs, yet one only is an active roller. The other either does not appear at all during this part of the work or, if it does, it merely clings passively to the pellet, and revolves in circles while its partner rolls. Thus it would at first sight seem that there was no cooperation between the two owners of the pellet in the effort to roll it along, but rather the very reverse, for the beetle clinging to the pellet not only makes no effort but rather acts as an obstruction to the free rolling of the smooth ball. But in reality this very obstruction is often to the advantage of the pair and I think it is to achieve this obstruction that the one beetle clings so firmly to the pellet and permits itself to be whirled round by its fellow. For when